When non-biologists
talk about biological evolution they often confuse two different aspects of the definition. On the one hand there is the question of whether or not modern
organisms have evolved from older ancestral organisms or whether modern
species are continuing to change over time. On the other hand there are questions
about the mechanism of the observed changes... how did evolution occur?
Biologists consider the existence of biological evolution to be a fact. It can be demonstrated today and the historical
evidence for its occurrence in the past is
overwhelming. However, biologists
readily admit that they are less certain of the exact mechanism of evolution; there
are several theories of the mechanism of
evolution.1
Before punctuated
equilibrium, most scientists assumed that evolutionary
change occurs slowly and continuously in almost all species, and that
new species originate either by slow divergence from parental stock of
sub-populations or by slow evolutionary transformation of the parental
stock
itself.2
Punctuated equilibrium
proposes that most species originate relatively suddenly
(over tens of thousands or hundreds of thousands of
years, rather than the millions of years assumed by
traditional theory) and then do not evolve significantly
for the rest of their time on Earth. Most species thus have a sudden or punctuated origin and then remain in stasis or equilibrium until extinction.2
Eldredge and Gould
proposed punctuated equilibrium to explain one of the
most notable features of the fossil record: most species seem to appear suddenly, already clearly differentiated from the earlier, similar species from which they
presumably evolved, and then remain unchanged until becoming extinct.2
Traditional
evolutionary theory proposed that gradual
evolutionary changes are rarely observed in the fossil record because that
record is radically incomplete. Fossils form only under
certain special conditions, fossil-bearing rocks are
eroded as well as deposited, and our knowledge even of
those fossils that have been formed is fragmentary.2
Eldredge and Gould
agree that the fossil record is incomplete, but contend that
it could not be incomplete enough to account for the near-complete absence of gradualistic change in the fossil record. Rather, they propose, species
normally originate too quickly for normal geological
processes to record the event.2